Dr. Y. Shibu Vardhanan

68818Vardhanan

Dr. . Y. Shibu Vardhanan
Professor

Department of Zoology, University of Calicut,
Calicut University PO, Malappuram District, Kerala 673635 India
Phone (Office): 0494-2407419
Mobile: +91- 9447108980
Email: This email address is being protected from spambots. You need JavaScript enabled to view it.; This email address is being protected from spambots. You need JavaScript enabled to view it.

Residence: C-5 Staff Quarters, University of Calicut, Calicut University, P.O, Malappuram District, Kerala, PIN: 673635, India

 

                                         Biography

Sl.

No.

Degree

Year

Subject

University/Institution

1

M.Sc.

1990 - 1992

Zoology

University College,

University of Kerala

 

2

 

Ph.D.

1994 - 1998

Toxicity Bioassays Histopathological studies & stress protein profile analysis

Aquatic Biology and Fisheries, University of Kerala

 

 

3

 

Post Doctoral work

 

2000 - 2004

 

Anaerobic biomethanation of polyphenols rich compounds

CSIR – National Institute for Interdisciplinary Science and Technology (NIIST),

Govt. of India Thiruvananthapuram, Kerala

 

Experience

Teaching: 21 years

Research: 25 years

Research Guidance

Ph.D.    Awarded – 6, Submitted - 1,  Ongoing - 6

M.Phil. Awarded – 9

Training

  • Advance training course equivalent to 4 European Credit Transfer System (4 ETS) in Geometric Morphometrics (GMM) Transmitting Science, S. L.U- NIF B65731903 -NIF- IVA ESB 65731903, C/Gardenia, 2- Urb. Can. Claramunt – 08784 Piera (Barcelona); Spain.
  • Eligible for procuring radioactive materials from Bhabha Atomic Research Centre (BARC), Mumbai and certification as Radiation Safety Officer (RSO)
  • Electron Microscope working experience, Training from All India Institutes of Medical Sciences (AIIMS), New Delhi.
  • Training on “Waste to Energy” from National Environmental Engineering Research Institute (NEERI), Nagpur, India.

Position held

  • Head of the department Zoology 2017-2019
  • Research Admission Committee (Subject expert) Kerala Forest Research Institute (KFRI), Peechi 2018 onwards
  • Research Admission Committee member Zoological Survey of India (ZSI), Calicut Research Admission Committee (Subject expert) Mahatma Gandhi University, 2018-2019 Research Admission Committee (Chairman) Mahatma Gandhi University, 2020
  • Director Centre of Health Science, University of Calicut 2011-2014 Member PG Board of studies, Bharathiar University

Current Research

  • Geometric morphometric: Shape and size to function

Morphological plasticity (inter and intra-population disparities) and adaptations are unsolvable using traditional taxonomic and molecular techniques; however, from these points, we can use advanced geometric morphometric methods to overcome the current obstacle in morpho-structural analysis. Our lab worked on innovative and inexpensive technology, which reliably demonstrated that shape and size differences. This information can be used to reveal the various morpho-function nature of an organism. Our research in this field focused on the multiple perspectives on organism’s development and adaptations, such as population-specific characteristic features, host-specific adaptions, geographical variations, niche preferences, and inter-and intra-specific competitions, pest surveillance and forecasting, morpho-structural evolutions, morphology trait-based phylogeny, morphological integration, phenotypic structural modular evolution, cryptic complex speciation, quantitative genetics. Our research has primarily focused on a broad, comparative study of insect morphological, structural variations, with a particular focus on wing and body architectural variations, as well as host-associated morphological adaptations of mites. Our research used several empirical and theoretical models and multivariate statistical approaches to analyze the morphological disparities. In addition, we correlated these morphological variations with the specific needs of species. So far, we have investigated the morphological differences of many insect taxa including, Lepidoptera, Hymenoptera, Diptera, Coleoptera, Hemiptera, and Odonata. In addition to the insect community, we analyzed the population structures of Bivalves and fishes with a particular emphasis on ecological conditions. Aside from the community level analysis, our research team discovered toxicity-specific morphological distortions and the effect of food on morphological developments with a focus on forensically important insects’ taxa.

  • Biomaterial characterization and their applications

Biomaterial engineering has enormous potential for the development and production of advanced functional and structural materials. Aquatic animal adhesive secretions (a protein- based natural biomaterial) can be utilized for various industrial and therapeutical applications. In a wet, saline, and highly turbulent environment, the unique nature of super hydrophilicity binds to a variety of solid surfaces. Therefore, biomimetic materials engineering has substantial potential for the development of advanced functional and structural materials. Ideally, these materials will be engineered using renewable resources and water-based processing strategies that closely mimic the green chemistry of biological systems. Our lab research focused on biomaterials from Bivalves; we used advanced, sophisticated instruments to characterize the biomaterials and a combination of biochemical and molecular techniques to understand the physicochemical properties of these biomaterials. Our research team is currently working on the Physico-chemical characterization of the byssus thread of native and invasive bivalves along the coastal belt of Kerala. We have already documented and sequenced the major adhesive proteins of three bivalve species, Perna Viridis, Perna Perna, and Mytilus strigata. We also studied and documented all mussel foot proteins' Physico-chemical structural and functional properties (78 mussel foot proteins from nine bivalve species, https://www.nature.com/articles/s41598-020- 59169-y). Besides, we are concentrating on industrial applications of mariculture waste such as byssus thread and Bivalve shells. Understanding the fundamental properties of these mussel- derived biomaterials helps us to create advanced ecofriendly biomimetic materials.

  • Toxicology

Quantitative assessment of developmental toxicity and risk assessment in insect model.

  • Nutrients immobilization

Nutrients immobilization is an advanced green strategy in nutrients administration in soil. Minerals from the natural sources are converted or augmented to a slow leaching complex, a future promising fertilizer, and resource management.

 

Patents Applied/Granted

  • Dr. Abhilash. P. L., & Dr. Y. Shibu Vardhanan. "A balanced nutrient cocktail for silkworms and a process of preparation thereof". University of Calicut, Kerala, Inda. Ref. No. TEM/E-1/85382/2024-CHE (Under process). 
  • P. P. Anand., & Dr. Y. Shibu Vardhanan. “A new cost-effective, long life, reusable filter cartridge for water treatment”. University of Calicut, Kerala, India. Ref. No: KSCSTE/143/2021-PF-PIC (Under process).

 

Links

ORCID iD: https://orcid.org/0000-0001-9820-886X

GOOGLE SCHOLAR: https://scholar.google.co.in/citations?user=F0A7l3oAAAAJ&hl=en&oi=ao

RESEARCH GATE: https://www.researchgate.net/profile/Shibu-Vardhanan-2

 

Projects

Ongoing 

  1. Principle Investigator SCIENCE & ENGINEERING RESEARCH BOARD (SERB) Research project entitled DEVELOPMENTOF ECO-FRIENDLYMUSSEL-INSPIRED FUNCTIONALIZED CATECHOL-RICH BIOMATERIALSAS REUSABLE ADSORBENT FOR WATER PURIFICATION SERB's sanction order No. EEQ/2021/000379 dated 04 March 2022, Total out lay 36.66 lakhs without project fellow (Ongoing project for 3 years).

Completed

  1. Coordinator UGC-SAP. “INTEGRATED AND INTERACTIVE DIGITALIZATION AND DNA BARCODING OF INSECTS AND MITES OF WESTERN GHATS”. 94.66Lakhs. Total Outlay Approved and Sanctioned = 60 Lakhs + 1 Project fellow (5 Year) Letter No. F.3-6/2012 (SAP-II) dated 10.10.2012.  (Completed)
  2. Coordinator UGC-BSR Project sanction letter. Lr. No.F.4-1/2006 (BSR)/7-363/2012(BSR) dated 4/7/2012. UGC-BSR Grant 20 lakhs + 5 Project fellow (5 Year) for infrastructure development (Completed)
  3. Principal Investigator (UGC-MRP) “Optimization of Bioreactor parameter for the Microbial expression of Bio-Adhesive foot protein from Indian Mussels Perna viridis and Perna perna”. Sanction No. F. No. 43-602/2014 (SR) 17-11-2015 Date of 25-01- 2016. Rs.13,3(Completed)
  4. Principal Investigator Minor research projects completed: Innovative Research Activities- wastewater characterization and management. 1) No. Lr. No. 149087/PLD-C- ASST-1/2014/Admn (1) dated: 27.1.2016. 2) U.O. No. 248/2016/Fin. dated 15/03/2016. Rs. 3,00,000.00 UGC XII plan. (Completed)

 

Recent publications (2018 onwards)

  1. Anzeer, F.M,, Santhosh, B., Anand, P. P., Shibu Vardhanan, Y., Aneesh, K. S., Gopalakrishnan, A., Krishnan, P. (2026) Highly Divergent Age-Specific Postembryonic Developmental Constraints and Phenotypic Plasticity in Hatchery-Reared Tropical Cloudy Damselfish, Dascyllus carneus Fischer, 1885. J Exp Zool A Ecol Integr Physiol. https://doi.org/10.1002/jez.70094.
  2. Mehabooba, B., Anand, P.P. & Shibu Vardhanan, Y. (2025). De novo transcriptome analysis of Perna perna L. (Bivalve) with functional and metabolic pathway analysis. Sci Data12, 1742. https://doi.org/10.1038/s41597-025-06028-y
  3. Paul, M. A., Anand, P. P., & Y. Shibu Vardhanan (2025). Electrophysiological cardiac response of housefly, Musca domestica Linnaeus, 1758 (Diptera: Muscidae) exposed to brief sublethal formalin vapor.  Toxicol. Environ. Health Sci. (2025). https://doi.org/10.1007/s13530-025-00261-z 
  4. Renu VV, & Y. Shibu Vardhanan. (2025). Transcriptomic insights into the effects of monocrotophos on Chrysomya megacephala: forensic implications of altered pupation time. Mol Biol Rep. 2025 Apr 9;52(1):378. doi: 10.1007/s11033-025-10455-w
  5. George, N., Mammili Kunnathu, A.L., Koodali Sherly, A. Joy. K., Kannan, A., Moosaripparambil, S. T., Y. Shibu Vardhanan., Meethal, V. K. (2025). Gut fungi and their interplay with other gut microbes play an important role in the larval development of the dengue vector Aedes albopictus. J Pest Sci. https://doi.org/10.1007/s10340-025-01916-z
  6. Seena, S., Anand, P. P., Y. Shibu Vardhanan. (2025). Comparative study of antennal sensilla of selected five genera of Buprestid beetles (Coleoptera: Buprestidae). Zoomorphology, 144(2). https://doi.org/10.1007/s00435-025-00709-3.
  7. Rahila, K., Y. Shibu Vardhanan. (2024). Comparative transcriptomic profiling of two pesticides, Acephate and Chlorantraniliprole in non-targeted insect model, Drosophila melanogaster. Pesticide Biochemistry & Physiology. 203, 106023. https://doi.org/10.1016/j.pestbp.2024.106023
  8. Chithra, B. K., Anand, P. P., Pushpalath, E., Y. Shibu Vardhanan., Mohammed Shareef, K. P. (2024). Bio-augmentative antibacterial activity of Pongamia pinnata (L.) gall extract with lemongrass oil formulation against selected Gram-negative bacteria. Current Science. 127(10): 1219-1226. 10.18520/cs/v127/i10/1219-1226
  9. Seena, S., Anand, P. P., Y. Shibu Vardhanan. (2023). Description of a new species of the genus Anthaxia (Haplanthaxia Reitter, 1911) from India with molecular barcoding and phylogenetic analysis. Journal of Threatened Taxa. 15(8): 23771-23777. https://doi.org/10.11609/jott.8178.15.8.23771-23777
  10. Seena, S., Anand, P. P., Y. Shibu Vardhanan. (2023). Morphology, ultrastructure and sexual dimorphism in atennal sensilla of Belionota prasina (Thunberg, 1789) (Coleoptera, Buprestidae). Journal of Insect Biodiversity and Systematics. 9(4): 663-678. https://doi.org/10.52547/jibs.9.4.663
  11. Meghana, R., Anand, P.P., Y. Shibu Vardhanan. (2023). Molecular and morphometric analyzes reveal host-specific cryptic speciation in a mite species, Tetranychus neocaledonicus (Andre, 1933) (Acari: Tetranychidae). Zootaxa. 5306, 1. https://doi.org/10.11646/ZOOTAXA.5306.1.3 
  12. Gayathri, M., Anand, P. P., & Y. Shibu Vardhanan. (2023). Visualization of early embryos of the wandering glider, Pantala flavescens (Fabricius, 1798) (Odonata: Libellulidae). Aquatic Insects. 44(4), 297-309. https://doi.org/10.1080/01650424.2023.2202182
  13. Gayathri, M., Anand, P. P., & Y. Shibu Vardhanan. (2023). Wing size, shape, and asymmetry analysis of the wandering glider, Pantala flavescens (Odonata: Libellulidae) revealed that hindwings are more asymmetric than the forewings. Biologia.78, 2763. https://doi.org/10.1007/s11756-023-01404-8
  14. Anand, P. P. & Y. Shibu Vardhanan. (2023). Molecular cloning, expression, mRNA secondary structure and immunological characterization of mussel foot proteins (Mfps) (Mollusca: Bivalvia). J. Biomolecular struct ure and Dynamics. https://doi.org/10.1080/07391102.2023.2166996
  15. Anand, P. P., Seena, S., Girish Kumar. P., & Shibu Vardhanan, Y. (2023). Species morphospace boundary revisited through wing phenotypic variations of Antodynerus species (Hymenoptera: Vespidae: Eumeninae) from the Indian continent. Front. Ecol. Evol. 10. https://doi.org/10.3389/fevo.2022.965577
  16. Shilpa, M. P., Anand, P.P., Y. Shibu Vardhanan., Manogem, E.M. (2022). High and lowland dependent wing phenotypic variation of the dark blue tiger butterfly, Tirumala septentrionis (Butler, 1874) (Lepidoptera: Nymphalidae) with FE-SEM wing scales nanomorphology. Zoologischer Anzeiger. 302(1). http://dx.doi.org/10.1016/j.jcz.2022.12.002.
  17. Shameema, K., Anand, P. P., Y. Shibu Vardhanan. (2022). Comparative dose-dependent stress responses of endosulfan and its isomers (α & β) in non-targeted insect model, Drosophila melanogaster Meign. Int. J. Tropical Insect Sci. 42: 3193-3202. https://doi.org/10.1007/s42690-022-00871-x.
  18. Anand, P.P., Mahima, K.V., Y. Shibu Vardhanan. (2022). Caste-specific quantitative genetics and phylogenetic signal analysis revealed the morphological adaptation of Asian weaver ant, Oecophylla smaragdina (Hymenoptera, Formicidae). J. Insect Biodiversity. 8(4): 595-615. http://dx.doi.org/10.52547/jibs.8.4.595.
  19. Anand, P.P., Seena, S., Jinsha Peter., Y. Shibu Vardhanan. (2022). Detection of geographical specific plasticity and the effect of natural selection pressure on the wing size and shape of Bactrocera dorsalis (Diptera: Tephritidae). Biologia, 77(4). http://dx.doi.org/10.1007/s11756-022-01059-x.
  20. Seena, S., Anand, P. P., Y. Shibu Vardhanan. (2022). Four new species from the Agrilus wittemani species-group (Coleoptera: Buprestidae: Agrilini) from Silent Valley National Park, Southern Western Ghats, Kerala, India. J. Asia-Pacific Entomology. 25(2): 101910. https://doi.org/10.1016/j.aspen.2022.101910
  21. Anand, P.P., & Shibu Vardhanan, Y. (2021). Dye and metal ion adsorption ability of Asian green mussel byssus thread complex; their microscopic and thermal property characterization. Environmental Technology. 44(3): 354-370. https://doi.org/10.1080/09593330.2021.1971776
  22. Mahima, K.V., Anand, P.P., Seena, S., Shameema, K., Manogem, E.M., & Y. Shibu Vardhanan. (2021). Caste-specific phenotypic plasticity of Asian weaver ants: Revealing the allometric and non-allometric component of female caste system of Oecophylla smaragdina (Hymenoptera: Formicidae) by using geometric morphometrics. Sociobiology; 68(2), e5941, eISSN: 2447-8067. Doi: 10.13102/sociobiology. v68i2.5941
  23. Anand, P.P., & Y. Shibu Vardhanan. (2021). Deleterious missense single nucleotide polymorphism alters the structural conformation of human anti-apoptotic Bcl-2 protein. Current Science. 20(4). Doi: 10.18520/cs/v120/i4/666-675.
  24. Aiswarya, D.Y., Shibu Vardhanan & Pushpalatha, E. (2021). Effect of xenobiotics on adult banana pseudo stem weevil Odoiporus longicollis oliv. (Coleoptera: Curculionidae), using landmark based geometric morphometric analysis. International Journal of Entomology Research. 6(1). ISSN 2455-4758
  25. Shameema, K., Anand, P.P., & Y. Shibu Vardhanan. (2021). Protective effect of Catharanthus roseus plant extracts against endosulfan and its isomers induced impacts on non-targeted insect model, Drosophila melanogaster and live brain cell imaging. Comparative Biochemistry and Physiology, Part C. 240(108916). https://doi.org/10.1016/j.cbp.2020.108916ISSN: 1532-0456
  26. Siva Parasad, M.S., Y. Shibu Vardhanan., Shaji Prabha, S.P., et al. (2021). MAOA-uVNTR polymorphism in male recidivist violent offenders in the Indian population. Archives of forensic medicine and Criminology. 2021. 70(4). Doi: 10.5114/amsik,2020.104863
  27. Shameema, K., Anand, P.P., & Y. Shibu Vardhanan. (2020). Evaluation of bioactive compounds present in the botanical Catharanthus roseus (L.) G. Don by GC-MS and FTIR spectroscopic analysis. International journal of Chemical studies. 8(5). Doi: https://doi.org/10.22271/ chemi.2020.v8.i5p.10447 ISSN 2349-8528
  28. Anand, P.P., & Y. Shibu Vardhanan. (2020). Computational modelling of wet adhesive mussel foot proteins (Bivalvia): Insights into the evolutionary convolution in diverse perspectives. Nature Scientific Report. 10(2612). Doi: https://doi.org/10.1038/s41598- 020-59169-y
  29. Aparana, T.P., Faizal, C.P., Y. Shibu Vardhanan et al.,(2020). Comparison of smear layer removal by MTAD, Tetraclean, Omix, NaOCl, coconut water, and saline as irrigating solution in primary teeth; an in vitro study. Journal of Dental; Research and Review. 7(3). Doi: 10.4103.jdrr.jdrr_39_20.
  30. Praveen Krishnakumar., Neethu.C.B., Sruthi, P., & Y. Shibu Vardhanan. (2019). Phytotoxicity and anaerobic sludge co-digestion inhibition of coconut husk Cocos nucifera (L.) (Palmae) leachate: Photo-oxidative remediation. Pollution Research. 38(2). ISSN: 0257- 8050
  31. Priya, M., Y. Shibu Vardhanan., Manilal, V.B. (2019). Response of anaerobic protozoa to oxygen tension in anaerobic system. International Microbiology. 22(3).Doi: 10.1007/s10123-019- 00058-4ISSN 1139-6709.
  32. Siva Prasad, M.S., & Y. Shibu Vardhanan. (2018). Evaluation of efficacy of collection techniques for human genomic DNA: MAOA-uVNTR polymorphism. International Journal of Scientific Research in biological science. 5(3).Doi: 10.26438/ijrbs/v5i3.611. ISSN: 2347-7520
  33. Shameema, K., & Y. Shibu Vardhanan. (2018). Insecticidal efficacy of aqueous and methanol extract of Catharanthus roseus (L.) G. Don. Against an insect model Drosophila melanogaster Meigen.. International Journal of Interdisciplinary research and innovations.6(3). ISSN 2348- 1218.
  34. Soumya, K., & Y. Shibu Vardhanan. (2018). Evaluation of toxic effects of Catharanthus roseus L., extract in Drosophila melanogaster M. with wing contour bioassay. Indian Journal of Entomology. 80(3).Doi: 10.5958/0974-8172.2018.002.05.5. ISSN 0367-8288
  35. Manogem, E.M., Soumya, P.K., & Y. Shibu Vardhanan. (2018). Geometric morphometrics approach on Genotoxicity evaluation of ecdyson agonist, Chromafenozide on Corcyra cephalonica Stainton (Lepidoptera: Pyralidae). International Journal of Engineering & technology. 7(4).
  36. Meharban, M.P., & Y. Shibu Vardhanan. (2018). Extraction of virgin coconut oil from the testa free albuminous endosperm through yeast mediated aqueous fermentation system: Fourier transform infrared spectra (FT-IR). International journal for research in Applied Science and Engineering. 6(3).Doi: 10.22214/ijraset.2018.3250.

Conferences (International/ National - Recent)

  1. Anand, P. P. & Y. Shibu Vardhanan. (2025). Functionalized catechol-rich biomaterial (mussel byssus thread) used as a reusable material for water purification. International conference on Advanced materials and Characterization – ICAMC 2025, Scheduled August 11-13, 2025 at Sathyabama Institute of Science and Technology, Chennai, India. Oral Presentation

  2. Anand, P. P. & Y. Shibu Vardhanan (2024). Sustainable usage of mussel byssus: A mariculture waste material as a reusable biomaterial for water purification. National conference on Marine Pollution & Ecotoxicology. Conducted by CUSAT, kochi, Kerala, from 25 to 27th of September 2024. *Secured the Best poster presentation award.  

  3. Mehabooba, B., & Y. Shibu Vardhanan. (2024). The effect of pH-Induced electrophysiological cardiac stress response analysis in Asian green mussel, Perna viridis L.  National conference on Marine Pollution & Ecotoxicology. Conducted by CUSAT, kochi, Kerala, from 25 to 27th of September 2024.  Poster presentation.

  4. Anand, P.P., & Shibu Vardhanan. Y. (2023). Identification, sequencing, and characterization of novel adhesive mussel foot proteins from Indian green mussel, Perna viridis (Linnaeus, 1758). International Conference on “Recent Advances in Animal Biology and Conservation”. Organized by Department of Zoology, University of College, Thiruvananthapuram, 19th December to 21st December, 2023. *Secured the Best oral presentation award.

  5. Anand, P.P., & Shibu Vardhanan. Y. (2023). Functionalized catechol-rich biomaterial (mussel byssus thread) used as a reusable material for water purification. National Conference on “Emerging Trends In Health Care Biotechnology: Innovations, Challenges and Future Prospects. Organized by Regional Cancer Center, Thiruvananthapuram, 30th November to 2nd December, 2023. Oral presentation.

  6. Anand, P.P., & Shibu Vardhanan. Y. (2023). Sustainable usage of byssus: A mariculture waste materials as a reusable biomaterial for water purification. International Seminar on “Ecotechnological Perspectives for sustainable development”. Organized by School of Bioscience, MG University, Kottayam. Oral presentation. 21st and 22nd March, 2023. Awarded: * Secured the Best oral presentation.
  7. Anand, P.P., & Shibu Vardhanan. Y. (2022). Dye and metal ion removal ability of byssus thread complex: A strategic value addition of mariculture waste. 34th Kerala Science Congress, Govt. of Kerala. India. Poster presentation.
  8. P. P. Anand., & Y. Shibu Vardhanan. (2021). Immunogenicity, allergenicity, antigenicity and expression analysis of wet adhesive mussel foot proteins of Asian green mussel, Perna viridis (Linnaeus, 1758).4th International conference on Biological and Biomimetic Adhesives. University of Aveiro, Portugal. Paper Presentation. International.
  9. P. P. Anand., Shameema K & Y. Shibu Vardhanan. (2021). In silico structural characterization of wet adhesive mussel foot proteins of Asian green mussel, Perna viridis (Linnaeus, 1758). 4th International conference on Biological and Biomimetic Adhesives. University of Aveiro, Portugal. Poster Presentation. International.

Workshops (International/ National - Recent)

  1. Next generation sequencing (NGS) – Data analysis. Unibiosys Biotech Research Lab, Kerala, India. National. 2019.
  2. Advances in Biomedical Mass Spectrometry and metabolomics. SAHA Institute of Nucear Physics Kolkata, India. Workshop. International. 2019.
  3. Advance training course equivalent to 4 European Credit Transfer System (4 ETS) in Geometric Morphometrics (GMM). Transmitting Science, S. L.U- NIF B65731903 - NIF- IVA ESB 65731903, C/Gardenia, 2- Urb. Can. Claramunt – 08784 Piera (Barcelona); Spain. Workshop: Training course. International. 2016

Life Member

  1. Indian Association of Nuclear Chemists and Allied Scientists (IANCAS) Bhabha Atomic Research Centre, Mumbai.
  2. Society of Fisheries Technologists (India).
  3. Kerala Academy of Science
  4. Ethological Society of India

 

 

 

 

 

 

 

 

 

 

Contact Us

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 Department of Zoology
University of Calicut
Calicut University PO
Malappuram Dist
PIN 673635
Kerala, India 

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